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The¡¡relation¡¡of¡¡the¡¡New¡¡Guinea¡¡fauna¡¡to¡¡that¡¡of¡¡Australia¡¡is¡¡very¡¡close¡£¡¡It¡¡is¡¡best¡¡marked¡¡in¡¡the¡¡Mammalia¡¡by¡¡the¡¡abundance¡¡of¡¡marsupials£»¡¡and¡¡the¡¡almost¡¡¡¡complete¡¡absence¡¡of¡¡all¡¡other¡¡terrestrial¡¡forms¡£¡¡In¡¡birds¡¡it¡¡is¡¡less¡¡striking£»¡¡although¡¡still¡¡very¡¡clear£»¡¡for¡¡all¡¡the¡¡remarkable¡¡old¡­world¡¡forms¡¡which¡¡are¡¡absent¡¡from¡¡the¡¡one¡¡are¡¡equally¡¡so¡¡from¡¡the¡¡other£»¡¡such¡¡as¡¡Pheasants£»¡¡Grouse£»¡¡Vultures£»¡¡and¡¡Woodpeckers£»¡¡while¡¡Cockatoos£»¡¡Broad¡­tailed¡¡Parrots£»¡¡Podargi£»¡¡and¡¡the¡¡great¡¡families¡¡of¡¡the¡¡Honeysuckers¡¡and¡¡Brush¡­turkeys£»¡¡with¡¡many¡¡others£»¡¡comprising¡¡no¡¡less¡¡than¡¡twenty¡­four¡¡genera¡¡of¡¡land¡­birds£»¡¡are¡¡common¡¡to¡¡both¡¡countries£»¡¡and¡¡are¡¡entirely¡¡confined¡¡to¡¡them¡£

When¡¡we¡¡consider¡¡the¡¡wonderful¡¡dissimilarity¡¡of¡¡the¡¡two¡¡regions¡¡in¡¡all¡¡those¡¡physical¡¡conditions¡¡which¡¡were¡¡once¡¡supposed¡¡to¡¡determine¡¡the¡¡forms¡¡of¡¡life¡­Australia£»¡¡with¡¡its¡¡open¡¡plains£»¡¡stony¡¡deserts£»¡¡dried¡¡up¡¡rivers£»¡¡and¡¡changeable¡¡temperate¡¡climate£»¡¡New¡¡Guinea£»¡¡with¡¡its¡¡luxuriant¡¡forests£»¡¡uniformly¡¡hot£»¡¡moist£»¡¡and¡¡evergreenthis¡¡great¡¡similarity¡¡in¡¡their¡¡productions¡¡is¡¡almost¡¡astounding£»¡¡and¡¡unmistakeably¡¡points¡¡to¡¡a¡¡common¡¡origin¡£¡¡The¡¡resemblance¡¡is¡¡not¡¡nearly¡¡so¡¡strongly¡¡marked¡¡in¡¡insects£»¡¡the¡¡reason¡¡obviously¡¡being£»¡¡that¡¡this¡¡class¡¡of¡¡animals¡¡are¡¡much¡¡more¡¡immediately¡¡dependent¡¡on¡¡vegetation¡¡and¡¡climate¡¡than¡¡are¡¡the¡¡more¡¡highly¡¡organized¡¡birds¡¡and¡¡Mammalia¡£¡¡Insects¡¡also¡¡have¡¡far¡¡more¡¡effective¡¡means¡¡of¡¡distribution£»¡¡and¡¡have¡¡spread¡¡widely¡¡into¡¡every¡¡district¡¡favourable¡¡to¡¡their¡¡development¡¡and¡¡increase¡£¡¡The¡¡giant¡¡Ornithopterae¡¡have¡¡thus¡¡spread¡¡from¡¡New¡¡Guinea¡¡over¡¡the¡¡whole¡¡Archipelago£»¡¡and¡¡as¡¡far¡¡as¡¡the¡¡base¡¡of¡¡the¡¡Himalayas£»¡¡while¡¡the¡¡elegant¡¡long¡­horned¡¡Anthribidae¡¡have¡¡spread¡¡in¡¡the¡¡opposite¡¡direction¡¡from¡¡Malacca¡¡to¡¡New¡¡Guinea£»¡¡but¡¡owing¡¡to¡¡unfavourable¡¡conditions¡¡have¡¡not¡¡been¡¡able¡¡to¡¡establish¡¡themselves¡¡in¡¡Australia¡£¡¡That¡¡country£»¡¡on¡¡the¡¡other¡¡hand£»¡¡has¡¡developed¡¡a¡¡variety¡¡of¡¡flower¡­haunting¡¡Chafers¡¡and¡¡Buprestidae£»¡¡and¡¡numbers¡¡of¡¡large¡¡and¡¡curious¡¡terrestrial¡¡Weevils£»¡¡scarcely¡¡any¡¡of¡¡which¡¡are¡¡adapted¡¡to¡¡the¡¡damp¡¡gloomy¡¡forests¡¡of¡¡New¡¡Guinea£»¡¡where¡¡entirely¡¡different¡¡forms¡¡are¡¡to¡¡be¡¡found¡£¡¡There¡¡are£»¡¡however£»¡¡some¡¡groups¡¡of¡¡insects£»¡¡constituting¡¡what¡¡appear¡¡to¡¡be¡¡the¡¡remains¡¡of¡¡the¡¡ancient¡¡population¡¡of¡¡the¡¡equatorial¡¡parts¡¡of¡¡the¡¡Australian¡¡region£»¡¡which¡¡are¡¡still¡¡almost¡¡entirely¡¡confined¡¡to¡¡it¡£¡¡Such¡¡are¡¡the¡¡interesting¡¡sub¡­family¡¡of¡¡Longicorn¡¡coleoptera¡¡Tmesisternitae£»¡¡one¡¡of¡¡the¡¡best¡­marked¡¡genera¡¡of¡¡Buprestidae¡¡Cyphogastra£»¡¡and¡¡the¡¡beautiful¡¡weevils¡¡forming¡¡the¡¡genus¡¡Eupholus¡£¡¡Among¡¡butterflies¡¡we¡¡have¡¡the¡¡genera¡¡Mynes£»¡¡Hypocista£»¡¡and¡¡Elodina£»¡¡and¡¡the¡¡curious¡¡eye¡­spotted¡¡Drusilla£»¡¡of¡¡which¡¡last¡¡a¡¡single¡¡species¡¡is¡¡found¡¡in¡¡Java£»¡¡but¡¡in¡¡no¡¡other¡¡of¡¡the¡¡western¡¡islands¡£

The¡¡facilities¡¡for¡¡the¡¡distribution¡¡of¡¡plants¡¡are¡¡still¡¡greater¡¡than¡¡they¡¡are¡¡for¡¡insects£»¡¡and¡¡it¡¡is¡¡the¡¡opinion¡¡of¡¡eminent¡¡botanists£»¡¡that¡¡no¡¡such¡¡clearly¡­defined¡¡regions¡¡pan¡¡be¡¡marked¡¡out¡¡in¡¡botany¡¡as¡¡in¡¡zoology¡£¡¡The¡¡causes¡¡which¡¡tend¡¡to¡¡diffusion¡¡are¡¡here¡¡most¡¡powerful£»¡¡and¡¡have¡¡led¡¡to¡¡such¡¡intermingling¡¡of¡¡the¡¡floras¡¡of¡¡adjacent¡¡regions¡¡that¡¡none¡¡but¡¡broad¡¡and¡¡general¡¡divisions¡¡can¡¡now¡¡be¡¡detected¡£¡¡These¡¡remarks¡¡have¡¡an¡¡important¡¡bearing¡¡on¡¡the¡¡problem¡¡of¡¡dividing¡¡the¡¡surface¡¡of¡¡the¡¡earth¡¡into¡¡great¡¡regions£»¡¡distinguished¡¡by¡¡the¡¡radical¡¡difference¡¡of¡¡their¡¡natural¡¡productions¡£¡¡Such¡¡difference¡¡we¡¡now¡¡know¡¡to¡¡be¡¡the¡¡direct¡¡result¡¡of¡¡long¡­continued¡¡separation¡¡by¡¡more¡¡or¡¡less¡¡impassable¡¡barriers£»¡¡and¡¡as¡¡wide¡¡oceans¡¡and¡¡great¡¡contrast£º¡¡of¡¡temperature¡¡are¡¡the¡¡most¡¡complete¡¡barriers¡¡to¡¡the¡¡dispersal¡¡of¡¡all¡¡terrestrial¡¡forms¡¡of¡¡life£»¡¡the¡¡primary¡¡divisions¡¡of¡¡the¡¡earth¡¡should¡¡in¡¡the¡¡main¡¡serve¡¡for¡¡all¡¡terrestrial¡¡organisms¡£¡¡However¡¡various¡¡may¡¡be¡¡the¡¡effects¡¡of¡¡climate£»¡¡however¡¡unequal¡¡the¡¡means¡¡of¡¡distribution£»¡¡these¡¡will¡¡never¡¡altogether¡¡obliterate¡¡the¡¡radical¡¡effects¡¡of¡¡long¡­continued¡¡isolation£»¡¡and¡¡it¡¡is¡¡my¡¡firm¡¡conviction£»¡¡that¡¡when¡¡the¡¡botany¡¡and¡¡the¡¡entomology¡¡of¡¡New¡¡Guinea¡¡and¡¡the¡¡surrounding¡¡islands¡¡become¡¡as¡¡well¡¡known¡¡as¡¡are¡¡their¡¡mammals¡¡and¡¡birds£»¡¡these¡¡departments¡¡of¡¡nature¡¡will¡¡also¡¡plainly¡¡indicate¡¡the¡¡radical¡¡distinctions¡¡of¡¡the¡¡Indo¡­Malayan¡¡and¡¡Austro¡­¡¡Malayan¡¡regions¡¡of¡¡the¡¡great¡¡Malay¡¡Archipelago¡£


CHAPTER¡¡XL¡£

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